Martin ZUSCHIN, Univ. Prof. Mag. Dr.

Room number: 2B410 Geocenter, UZA II
Tel.:     +43-1-4277-53555
Fax:     +43-1-4277-9535

Research interests:

    Paleodiversity and faunal gradients in the the Cenozoic marine fossil record of Europe
    Palaeoecology, taphonomy and actuopalaeontology of benthic invertebrates
    Shell beds and skeletal concentrations
    Marine anoxia

Current funded projects:

    Historical ecology of the northern Adriatic Sea
    Conservation palaeobiology of oil-polluted tropical marine biota in the Arabian Gulf
    Biodiversity of Cenozoic molluscan faunas of Europe


    Current semester
    Previous semester



Zuschin, M. & Ebner, C. in press, Actuopaleontological characterization and molluscan biodiversity of a protected tidal flat and shallow subtidal at the northern Red Sea. Facies.

Zuschin, M. & Ebner, C. in press, Compositional fidelity of death assemblages from a coral reef-associated tidal flat and shallow subtidal lagoon in the northern Red Sea. Palaios.


Tomašových, A., Dominici, S,  Zuschin,M.,  Merle, D., 2014, Onshore–offshore gradient in metacommunity turnover emerges onlyover macroevolutionary time-scales. Proceedings of the Royal Society of London 281, 20141533. pdf

Riedel B., Pados, T., Pretterebner, K., Schiemer, L., Steckbauer, A., Zuschin, M., Stachowitsch M. 2014, Effect of hypoxia and anoxia on invertebrate behaviour: ecological perspectives from species to community level. Biogeosciences 11, 1491-1518. pdf

Langlet, D., Baal, C., Geslin, E., Metzger, E., Zuschin, M., Riedel, B., Risgaard-Petersen, N., Stachowitsch, M. and Jorissen, F. J., 2014, Foraminiferal specific responses to experimentally induced anoxia in the Adriatic Sea. Biogeosciences 11, 1775-1797. pdf

Zuschin, M., Harzhauser, M. Hengst, B., Mandic, O., Roetzel, R. 2014, Long-term ecosystem stability in a Lower Miocene estuarine succession. Geology  42,  p.1-4.  pdf

Chattopadhyay, D., Zuschin, M., Tomašových, A., 2014, Effects of a high-risk environment on  edge-drilling behavior: inference from Recent bivalves from Red Sea. Palaeobiology 40, 34-49. pdf


Blasnig, M., Riedel, B., Zuschin, M., Schiemer, L. and Stachowitsch, M, 2013, Short-term post-mortality predation and scavenging and longer-term recovery after anoxia in the northern Adriatic Sea, Biogeosciences 10, 7647–7659. pdf

Langlet, D., Geslin, E., Baal, C., Metzger, E., Lejzerowicz, F., Riedel, B., Zuschin, M., Pawlowski, J., Stachowitsch, M. and Jorissen, F. J., 2013: Foraminiferal survival after one year of experimentally induced anoxia, Biogeosciences 10, 7463-7480. pdf

Brand, U., Azmy, K. Bitner, M.A., Logan, A., Zuschin, M., Came, R. Ruggiero, E. 2013, Oxygen isotopes and MgCO3 in brachiopod calcite and a new paleotemperature equation. Chemical Geology 359, 23-31. pdf

Weber, K, Zuschin, M. 2013, Delta-associated molluscan life and death assemblages in the northern Adriatic Sea: Implications for palaeoecology, regional diversity and conservation. Palaeogeography, Palaeoclimatology, Palaeoecology 370, 77-91. pdf

Schneider, S, Harzhauser, M., Kroh, A., Lukeneder, A. & Zuschin, M., 2013, Ernstbrunn Limestone and Klentnice Beds (Kimmeridgian-Berriasian; Waschberg-Ždánice Unit; NE Austria and SE Czech Republic): state of the art and bibliography. Bulletin of Geosciences 88. 105-130. pdf

Kleemann, K. & Zuschin, M. 2013, Traces of predation on Recent terrestrial gastropod shells of the family Helicidae. Conchylia 43, 41-44.  pdf


Harzhauser, M., Neubauer, T., Mandic, O., Zuschin, M., 2012, A Middle Miocene endemic freshwater mollusc assemblage from an intramontane Alpine lake (Aflenz Basin, Eastern Alps, Austria). Paläontologische Zeitschrift 86, 23-41 pdf

Pretterebner, K., Riedel, B., Zuschin, M., Stachowitsch, M. 2012, Hermit crabs and their symbionts: reactions to artificially induced anoxia on a sublittoral sediment bottom. Journal of Experimental Marine Biology and Ecology 411, 23-33. pdf

Riedel B., Zuschin M., Stachowitsch M. 2012, Tolerance of benthic macrofauna to hypoxia and anoxia in shallow coastal seas: a realistic scenario. Marine Ecology-Progress Series 458, 39-52. pdf

Kroh, A. & Zuschin, M. 2012, Was bleibt vom Riff? Das Naturhistorische 9/2012, p. 79


Zuschin, M., Harzhauser, M. & Mandic, O., 2011, Disentangling palaeodiversity signals from a biased sedimentary record: An example from the Early to Middle Miocene of Central Paratethys Sea. In: McGowan, A. & Smith, A.B. (eds) Comparing the Geological and Fossil Records: Implications for Biodiversity Studies. Geological Society, London, Special Publications, 358, 123-139. pdf

Lukeneder S., Zuschin M., Harzhauser M., Mandic O., 2011: Spatiotemporal signals and palaeoenvironments of endemic molluscan assemblages in the marine system of the Sarmatian Paratethys. Acta Palaeontologica Polonica 56, 767-784. pdf

Stachowitsch, M., Riedel, B. Zuschin, M., 2011, The return of shallow shelf seas as extreme environments: anoxia and macrofauna reactions in the Northern Adriatic Sea. In: Altenbach, A.V., Bernhard, J.M. and Seckbach, J. (eds). Anoxia: Anoxia: Evidence for Eukaryote Survival and Paleontological Strategies. Book Series: Cellular Origin, Life in Extreme Habitats and Astrobiology (COLE) vol 21, pp353-368. Springer-Verlag. pdf

Zuschin, M. 2011, Book review of "Marine Macroecology" by J.D. Witman & K. Roy (Eds): Marine Ecology: an evolutionary perspective 32, 256-257. pdf

Janssen, R., Zuschin, M. Baal, Ch., 2011, Gastropods and their habitats from the northern Red Sea (Egypt: Safaga) Part 2: Caenogastropoda: Sorbeoconcha and Littorinimorpha. Annalen des Naturhistorischen Museums Wien 113A, 373-509. pdf

Sawyer, J.A, & Zuschin, M. 2011. Drilling predation in molluscs from the Early and Middle Miocene of the Central Paratethys. Palaios 26, 284-297. pdf

Kern, A., Harzhauser, M., Mandic, O., Roetzel, R., Coric, S., Bruch, A., Zuschin, M. 2011. Millenium-scale vegetation dynamics in an estuary at the onset of the Miocene climate optimum. Palaeogeography, Palaeoclimatology, Palaeoecology 304, 247-261. pdf


Haselmair A., Stachowitsch M., Zuschin, M., Riedel B. 2010. Behaviour and mortality of benthic crustaceans in response to experimentally induced hypoxia and anoxia in situ. Marine Ecology-Progress Series 414, 195-208. pdf

Titschack, J., Zuschin, M., Spötl C. Baal. C. 2010. The giant oyster Hyotissa hyotis from the northern Red Sea as a decadal-scale archive for seasonal environmental fluctuations in coral reef habitats. Coral Reefs 29, 1061-1075. pdf

Schoepf, V., Herler, J., Zuschin, M. 2010, Microhabitat use and prey selection of the coral-feeding snail Drupella cornus in the northern Red Sea. Hydrobiologia 641, 45-57. pdf

Sawyer, J.A, Zuschin, M. 2010, Intensities of drilling predation from molluscan assemblages along a transect through the northern Gulf of Trieste ( Adriatic Sea ). Palaeogeography, Palaeoclimatology, Palaeoecology 285, 152-173. pdf

Wagreich, M. & Zuschin, M. 2010. Sedimentology in Austria. IAS (International Association of Sedimentologists) Newsletter 229, p. 16-20.


Tomasovych, A, Zuschin, M. 2009, Variation in brachiopod preservation along a subtropical carbonate shelf-basin transect (Red Sea and Gulf of Aden ): Untangling the effects of shell specific and environmental factors. Palaios 24, 697-716. pdf

Zuschin, M., Mayrhofer, S. 2009. Brachiopods from cryptic coral reef habitats in the Northern Red Sea. Facies 55, 335-344. pdf

Zuschin, M., Gebhardt, H. 2009 Octocorals as hosts to serpulid-macroids from the Cretaceous of the Potiguar Basin (NE Brazil).  Lethaia 42, 381-382. pdf

Zuschin, M., Janssen, R., Baal, Ch., 2009, Gastropods and their habitats from the northern Red Sea (Egypt: Safaga) part 1: Patellogastropoda, Vetigastropoda and Cycloneritimorpha. Annalen des Naturhistorischen Museums Wien 111A: 74-158, 19 plates. pdf

Sawyer, J.A., Zuschin, M., Riedel, B., Stachowitsch, M. 2009. Predator-prey interactions from in situ time-lapse observations of a sublittoral mussel bed in the Gulf of Trieste (Northern Adriatic). Journal of Experimental Marine Biology and Ecology 371, 10-19. pdf

Zuschin M. & Stachowitsch M., 2009, Epifauna-dominated benthic shelf assemblages: lessons from the modern Adriatic Sea. Palaios 24, 148-158. pdf

Zuschin, M. 2009, Book Review of “Phylogeny and Evolution of the Mollusca” by Ponder, W.F. & Lindberg, D.R. Marine Ecology: an evolutionary perspective 30, 269. pdf

Zuschin, M. & Hofmann, T. (eds.) 2009, 15. Jahrestagung der Österreichischen Paläontologischen Gesellschaft, 9. – 10- Oktober 2010, Tagungsprogramm, Abstracts, Exkursion. Berichte der Geologischen Bundesanstalt 81, 49 p. pdf


Harzhauser, M., Kern, A., Soliman, A., Minati, K, Piller, W.E., Danielopol, D., Zuschin, M. 2008. Centennial- to decadal-scale environmental shifts in and around Lake Pannon (Vienna Basin) related to a major Late Miocene lake-level rise. Palaeogeography, Palaeoclimatology, Palaeoecology 270, 202-215. pdf

Riedel B., Zuschin M., Stachowitsch M. 2008. Dead zones: a future worst-case scenario for Northern Adriatic biodiversity. In: Climate warming and related changes in Mediterranean marine biota. F. Briand (ed.). CIESM Workshop Monograps 35: 73-77. pdf

Riedel B., Zuschin M., Haselmair A., Stachowitsch M., 2008,. Oxygen depletion under glass: behavioural responses of benthic macrofauna to induced anoxia in the Northern Adriatic. Journal of Exp Marine Biology and Ecology 367, 17-27. pdf

Riedel, B., Stachowitsch, M, Zuschin, M. 2008, Sea anemones and brittle stars: unexpected predatory interactions during induced in situ oxygen crises. Marine Biology 153, 1075-1085, pdf

Logan, A., Tomasovych, A., Zuschin, M., Grill, B., 2008, Recent brachiopods from the Red Sea and Gulf of Aden. Fossils and Strata 54, 299-309. pdf

Zuschin, M. & Stachowitsch, M. 2008, Book review of "The northern Adriatic ecosystem: deep time in a shallow sea" by Frank K. McKinney: Marine Ecology: an evolutionary perspective 29, 411-412. pdf

Zuschin, M. & Hofmann, T. (eds.) 2008, 14. Jahrestagung der Österreichischen Paläontologischen Gesellschaft, 15.-18. Mai 2008, Tagungsprogramm, Abstracts, Exkursion. Berichte der Geologischen Bundesanstalt 75, 59 p. pdf


Stachowitsch M., Riedel, B, Zuschin, M., Machan, R. 2007, Oxygen depletion and benthic mortalities: the first in situ experimental approach to documenting an elusive phenomenon. Limnology & Oceanography: Methods 5, 344-352. pdf

Zuschin, M. 2007b, Review of "The oceans and climate", by Grant Bigg. Palaios 22. pdf

Zuschin, M. 2007a, Review of "Echinoidea neogenica", by Andreas Kroh. Lethaia 40, p. 199. pdf

Dominici, S. & Zuschin, M. 2007. Sea-level change and the structure of marine ecosystems. Palaios 22, 225-227. pdf

Zuschin, M., Baal, Ch., 2007, Large gryphaeid oysters as habitats for numerous sclerobionts: a case study from the northern Red Sea. Facies 53, 319-327. pdf

Zuschin, M. Stachowitsch, M. 2007, The distribution of molluscan assemblages and their postmortem fate on coral reefs in the Gulf of Aqaba (northern Red Sea). Marine Biology 151, 2217-2230. pdf

Zuschin, M., Harzhauser, M. and Mandic, O. 2007. The stratigraphic framework of fine-scale gradual and disjunct faunal replacements in the Middle Miocene of the Vienna basin (Austria). Palaios 22, 286-297. pdf

Zuschin M. 2007. Evolutionäre Paläoökologie mariner Lebensräume. In: Gusenleitner, F., Aubrecht, G., Aescht, E., Pfosser, M. (eds). Evolution-Phänomen Leben. Denisia 20, p. 139-148. pdf


Zuschin, M., Harzhauser, M. and Sauermoser, K. 2006. Patchiness of local species richness and its implication for large-scale diversity patterns: an example from the middle Miocene of the Paratethys. Lethaia 39, 65-78. pdf

Harzhauser, M., Wanzenböck, G. & Zuschin, M. (eds.) 2006, 12. Jahrestagung der Österreichischen Paläontologischen Gesellschaft, Tagungsprogramm, Abstracts, Exkursion. Eigenverlag, Naturhistorisches Museum Wien, 64 pages, pdf

Zuschin, M. 2006. Review of: "Bivalvia neogenica (Soenoidea-Clavagelloidea)" di Schultz Ortwin. - Notiziario S.I.M., Pubblicazione quadrimestrale della Società Italiana di Malacologia, 24/1-4: 29/Recensioni. - Milano.

Zuschin, M. & Harzhauser, M. 2006. Report on the “The 12th Congress R.C.M.N.S.”. IAS (International Association of Sedimentologists) Newsletter 202, p. 8-9.


Zuschin, M. & Oliver, P.G. 2005. Diversity patterns of bivalves in a coral dominated shallow-water bay in the northern Red Sea – high species richness on a local scale. Marine Biology Research 1, 396-410. pdf

Taylor, J.D., Glover, E.M., Zuschin, M., Dworschak, P.C., Waitzbauer, W. 2005. Another bivalve with dreadlocks: living Rasta lamyi from Aqaba, Red Sea (Bivalvia: Lucinidae). Journal of Conchology 38, 489-497.

Dominici, S. & Zuschin, M. 2005. Infidelities of fossil assemblages. Lethaia 39, 381-382. pdf

Zuschin, M., M. Harzhauser and O. Mandic. 2005. Influence of size-sorting on diversity estimates from tempestitic shell beds in the middle Miocene of Austria.- Palaios 20, 142-158. pdf


Coric, S., Harzhauser, M., Hohenegger, J., Mandic, O., Pervesler, P., Roetzel, R., Rögl, F., Scholger, R., Spezzaferri, S., Stingl, K., Svabenicka, L., Zorn, I. & Zuschin, M. 2004. Stratigraphy and correlation of the Grund Formation in the Molasse Basin, northeastern Austria (Middle Miocene, Lower Badenian): Geologica Carpathica 55, 207-215. pdf

Pervesler, P. & Zuschin, M. 2004. A lucinoid bivalve trace fossil Saronichnus abeli igen. et isp. nov. from the Miocene molasse deposits of Lower Austria, and its environmental significance. Geologica Carpathica 55, 111-115. pdf

Zuschin, M., Harzhauser, M. & Mandic, O. 2004. Spatial variability within a single parautochthonous Paratethyan tidal flat deposit (Karpatian, Lower Miocene – Kleinebersdorf, Lower Austria): Courier Forschungsinstitut Senckenberg 246, 153-168. pdf

Zuschin, M., Harzhauser, M. & Mandic, O. 2004. Taphonomy and Palaeoecology of the Lower Badenian (Middle Miocene) molluscan assemblages at Grund (Lower Austria): Geologica Carpathica 55, 117-128. pdf


Zuschin, M. & Oliver, P.G. 2003. Bivalves and bivalve habitats in the northern Red Sea. Verlag des Naturhistorischen Museums, Wien, 304p, 57plates.  

Harzhauser, M., Mandic, O., Zuschin, M. 2003. Changes in Paratethyan marine molluscs at the Early/Middle Miocene transition: diversity, paleogeography and paleoclimate. – Acta Geologica Polonica 53, 323-339. pdf  

Zuschin, M. & Oliver, P.G. 2003. Fidelity of molluscan life and death assemblages on sublittoral hard substrata around granitic islands of the Seychelles. Lethaia 36, 133-150. pdf

Zuschin, M., Stachowitsch, M. & Stanton, R.J., Jr. 2003. Patterns and processes of shell fragmentation in modern and ancient marine environments. Earth-Science Reviews 63, 33-82. pdf

Zuschin, M. 2003. Report on the “First Austrian Reef Workshop”. IAS (International Association of Sedimentologists) Newsletter 188, p. 9-10.


Zuschin, M. & Stanton, R.J., Jr. 2002. Paleocommunity Reconstruction from Shell Beds - A Case Study from the Main Glauconite Bed, Eocene, Texas. Palaios 17, 602-614. pdf

Mandic, O.; Harzhauser, M.; Spezzaferri, S. & Zuschin, M. 2002. The paleoenvironment of an early Middle Miocene Paratethys sequence in NE Austria with special emphasis on paleoecology of mollusks and foraminifera. - Geobios, Mémoire spécial 24, 193-206.  pdf

Zuschin, M. 2002. Review of: Bivalvia neogenica (Nuculacea-Unionacea), by Ortwin Schultz, 2001, In Catalogus Fossilium Austriae, Vol 1, Part 1. Verlag der Österreichischen Akademie der Wissenschaften, Wien. Lethaia 35, p. 278


Zuschin, M., Mandic, O., Harzhauser, M. & Pervesler, P. 2001. Fossil evidence for chemoautotrophic bacterial symbiosis in the thyasirid bivalve Thyasira michelottii from the Middle Miocene (Badenium) of Austria.- Historical Biology 15: 223-234.

Zuschin, M. & Stanton, R.J., Jr. 2001. Experimental measurement of shell strength and its taphonomic interpretation.- Palaios 16, 161-170. pdf

Zuschin, M., Hohenegger, H., Steininger, F.F. 2001. Molluscan assemblages on coral reefs and associated hard substrata in the northern Red Sea.- Coral Reefs 20, 107-116.pdf

Grill, B. & Zuschin, M. 2001. Modern shallow- to deep-water bivalve death assemblages in the Red Sea – ecology and biogeography.- Palaeogeography, Palaeoclimatology, Palaeoecology 168: 75-96. pdf

Oliver, P.G., & Zuschin, M. 2001. Minute Veneridae and Kelliellidae from the Red and Arabian Seas with a redescription of Kellia miliacea Issel, 1869.- Journal of Conchology, 37(2): 213-230. pdf

Harzhauser, M., Mandic, O., Zuschin, M., Piller, W. Hofmann, T. 2001. From the Paratethys Sea to Lake Pannon. Vienna Basin and Eisenstradt-Sopron Basin. Field guide World Congress of Malacology 2001, Vienna, Austria. 16 pages, 4 plates.


Zuschin, M., Hohenegger, H., Steininger, F.F. 2000. A comparison of living and dead molluscs on coral reef associated hard substrata in the northern Red Sea – implications for the fossil record.- Palaeogeography, Palaeoclimatology, Palaeoecology 159: 167-190. pdf

Oliver, P.G., & Zuschin, M. 2000. Additions to the bivalve fauna of the Red Sea with descriptions of new species of Limopsidae, Limidae, Semelidae, and Veneridae.- Journal of Conchology, 37(1), 17-38.  pdf


Zuschin, M., Stachowitsch, M. Pervesler, P & Kollmann, H. 1999. Structural features and taphonomic pathways of a high-biomass epifauna in the northern Gulf of Trieste, Adriatic Sea.- Lethaia 32, 299-317. pdf

Schlaf, J., Zuschin, M. & Piller, W.E. 1999. Origin and paleoenvironment of thick coquina accumulations on a Lower Norian (Upper Triassic) carbonate slope (Julian Alps, Slovenia).- Zentralblatt für Geologie und Paläontologie, Teil I, 1997 (7-9): 1153-1166.

Roetzel, R., Pervesler, P., Daxner-Höck, G., Harzhauser, M., Mandic, O., Zuschin, M.  and Cicha, I. 1999. C4 Grund - Kellergasse. Field guide Arbeitstagung der Geologischen Bundesanstalt. Retz – Hollabrunn: 328-334.


Zuschin, M. & Hohenegger, J., 1998. Subtropical coral-reef associated sedimentary facies characterized by molluscs, Northern Bay of Safaga, Red Sea, Egypt.- Facies 38, 229-254. pdf


Schlaf, J. & Zuschin, M., 1997. Sediment compaction as formational process for nodular limestones of the Anisian (Middle Triassic) Han Bulog Formation (Hydra; Greece).- Zentralblatt für Geologie und Paläontologie, Teil I, 1996 (5/6): 609-618.

Zuschin, M. & Piller, W.E., 1997. Molluscan Hard-Substrate Associations in the Northern Red Sea.- PSZNI: Marine Ecology, 18, 361-378

Zuschin, M. & Piller, W.E., 1997. Gastropod shells recycled - an example from a rocky tidal flat in the northern Red Sea.- Lethaia, 30, 127-134.pdf

Zuschin, M. & Piller, W.E., 1997. Bivalve distribution on coral carpets in the northern Bay of Safaga (Red Sea, Egypt) and its relation to environmental parameters.- Facies, 37, 193-204. pdf


Zuschin, M. & Pervesler, P., 1996 Secondary hardground-communities in the Northern Gulf of Trieste, Adriatic Sea.- Senckenbergiana marit., 28(1/3): 53-63

Zuschin, M., 1996. Influence of autochthonous biogenic material on textural characteristics of fine-grained terrigenous sediments.- Zentralblatt für Geologie und Paläontologie Teil I, 1995 (1/2): 387-398.


Zuschin, M. &. Piller, W.E., 1994. Sedimentology and facies zonation along a transect through the Gulf of Trieste, Northern Adriatic Sea.- Beiträge zur Paläontologie, 18:75-114. pdf